Class 11 : Biology (In English) – Lesson 17: Locomotion and Movement
EXPLANATION & SUMMARY
π±β¨ Introduction
π§ Movement is one of the fundamental characteristics of living organisms. It refers to a change in body position or a part of the body.
βοΈ Locomotion is a special type of movement that results in a change of location of the organism β like walking, running, flying, or swimming.
πΏ In plants, movement is usually growth-oriented, while in animals, locomotion helps in finding food, shelter, mates, and escaping predators.
π‘ Concept:
Movement = any displacement (can be internal or external)
Locomotion = displacement of the entire body
All locomotions are movements, but not all movements are locomotions
𧬠Types of Movement
Organisms exhibit three major types of movements:
1οΈβ£ Amoeboid Movement
Seen in amoeba, WBCs
Achieved by pseudopodia (cytoplasmic extensions)
Involves cytoplasmic streaming and actin filament rearrangement
βοΈ Example: Movement of macrophages during phagocytosis.
2οΈβ£ Ciliary Movement
Uses tiny hair-like cilia
Found in paramecium, tracheal epithelium, fallopian tubes
Coordinated beating moves fluids or the entire organism
π‘ In humans:
β‘οΈ Respiratory tract β clears mucus
β‘οΈ Oviduct β moves ovum toward uterus
3οΈβ£ Muscular Movement
Involves contraction and relaxation of muscle fibers
Responsible for locomotion in higher animals
Works under neural control
βοΈ Seen in limbs, heart, alimentary canal.
𦴠Human Muscular System
π§ Muscles are specialized contractile tissues derived from mesoderm.
π Classified into three main types:

π’ 1οΈβ£ Skeletal (Striated / Voluntary) Muscles
Attached to bones via tendons
Multinucleated, striated (light and dark bands)
Voluntary control
Function: locomotion, posture, body movement
πͺ Examples: Biceps, triceps, quadriceps

π΅ 2οΈβ£ Smooth (Unstriated / Involuntary) Muscles
Found in walls of visceral organs (stomach, intestine, blood vessels)
Uninucleated, non-striated
Involuntary control
Function: movement of food, vessel constriction

π΄ 3οΈβ£ Cardiac Muscles
Found only in heart β€οΈ
Striated, branched, uninucleated
Involuntary, rhythmic contraction
Function: pumps blood continuously
π‘ Concept: All muscle contractions are powered by ATP and regulated by calcium ions (CaΒ²βΊ).

βοΈ Structure of Skeletal Muscle

𧬠Muscle β bundles of muscle fascicles β muscle fibers (cells) β myofibrils
Each myofibril shows alternating dark (A band) and light (I band), giving striated appearance.
π Sarcomere = structural & functional unit of contraction
Bound by Z-lines
πΉ Components of a Sarcomere

Band Description
A band Dark band (myosin + overlap with actin)
I band Light band (only actin)
H zone Center of A band (only myosin)
M line Midline of sarcomere
Z line Boundary between sarcomeres
π‘ During contraction:
Sarcomere length decreases
I band & H zone shorten, A band constant
π§ Sliding Filament Theory
π§ͺ Proposed by Huxley and Niedergerke
βοΈ Explains muscle contraction as sliding of actin filaments over myosin filaments.
π Steps:
1οΈβ£ Impulse from motor neuron β release of CaΒ²βΊ from sarcoplasmic reticulum
2οΈβ£ CaΒ²βΊ binds troponin, exposes binding sites on actin
3οΈβ£ Myosin head attaches to actin β forms cross bridge
4οΈβ£ ATP hydrolysis β myosin pulls actin inward (power stroke)
5οΈβ£ New ATP breaks cross bridge β cycle repeats
π‘ Energy: ATP
π‘ Ions: CaΒ²βΊ, MgΒ²βΊ
βοΈ Result: Shortening of sarcomere = contraction.
𧬠Neuromuscular Junction

π Synapse between motor neuron and muscle fiber
π§ Impulse β releases acetylcholine (ACh) into synaptic cleft
β‘οΈ Triggers depolarization β contraction
βοΈ AChE (acetylcholinesterase) breaks down ACh to stop contraction.
πͺ Types of Muscle Contractions

1οΈβ£ Isotonic β muscle shortens, movement occurs
2οΈβ£ Isometric β tension changes, length constant
3οΈβ£ Tetanic β sustained contraction from rapid stimuli
βοΈ Example: Holding weight β isometric; lifting β isotonic.
𦴠Skeletal System
𦴠Framework of bones and cartilage that supports body and aids movement.
π Human skeleton:
Axial skeleton β skull, vertebral column, ribs, sternum
Appendicular skeleton β limbs and girdles
π§ Total bones: 206 in adult

πΏ Axial Skeleton (80 bones)



Skull: 22 bones (8 cranial + 14 facial)
Vertebral column: 26 vertebrae
Ribs: 12 pairs
Sternum: 1
π‘ Protects brain, spinal cord, heart, lungs.
πͺ Appendicular Skeleton (126 bones)
Pectoral girdle: 4
Pelvic girdle: 2
Limbs: 60 (upper) + 60 (lower)
βοΈ Enables movement and locomotion.
𦡠Limb Bones
ποΈ Upper Limb
Humerus β Radius + Ulna β Carpals β Metacarpals β Phalanges
π¦Ά Lower Limb
Femur β Tibia + Fibula β Tarsals β Metatarsals β Phalanges
βοΈ Longest bone: Femur
π‘ Strongest joint: Knee joint

π© Joints
π Articulation = point of contact between bones
Types (Structural basis):
1οΈβ£ Fibrous (Immovable) β skull sutures
2οΈβ£ Cartilaginous (Partially movable) β vertebral joints
3οΈβ£ Synovial (Freely movable) β limbs (shoulder, knee)
π§ͺ Synovial joint features:
Synovial fluid (lubrication)
Capsule, ligaments
Articular cartilage
π‘ Examples:
Ball & socket: shoulder

Hinge: elbow, knee

Pivot: neck (atlas-axis)
Gliding: wrist

Saddle: thumb
Condyloid: fingers
βοΈ Disorders of Muscles and Bones
1οΈβ£ Myasthenia gravis β autoimmune; blocks ACh receptors β weakness
2οΈβ£ Muscular dystrophy β genetic; degeneration of muscles

3οΈβ£ Tetanus β sustained contraction due to toxin
4οΈβ£ Osteoporosis β bone thinning (β calcium, estrogen)
5οΈβ£ Arthritis β inflammation of joints
6οΈβ£ Gout β uric acid accumulation in joints
7οΈβ£ Fracture β bone breakage
π‘ Proper nutrition, exercise, and posture maintain skeletal health.

π Importance of Locomotion
π§ Enables interaction with environment
β‘ Supports feeding, reproduction, protection
𧬠Maintains organism survival
πΏ In humans, essential for daily activities, sports, and circulation (muscle pump effect)
π Why This Lesson Matters
πͺ Explains how the body moves and mechanics of muscles
π§ Connects anatomy and physiology
βοΈ Foundation for sports medicine, orthopedics, neurobiology
π Highlights importance of calcium, exercise, and posture
π Quick Recap
π§ Movement = body part displacement
π Locomotion = movement + location change
πͺ Muscles: skeletal, smooth, cardiac
βοΈ Sarcomere = unit of contraction
𧬠Sliding filament theory: actin slides over myosin
π§ CaΒ²βΊ + ATP essential for contraction
𦴠Skeleton: 206 bones; axial + appendicular
π© Joints: fibrous, cartilaginous, synovial
β οΈ Disorders: Myasthenia, osteoporosis, arthritis
π Movement vital for life and activity
π Summary
Locomotion and movement form the basis of animal activity. Movements can be amoeboid, ciliary, or muscular. In humans, skeletal muscles enable locomotion through contraction based on the sliding filament theory. The skeletal system, with bones and joints, provides structure, support, and leverage. The nervous system and ions like calcium coordinate contraction. Joints permit various movements, while disorders such as arthritis and osteoporosis impair mobility. This system ensures organisms interact with their environment, sustain posture, and perform essential functions.
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QUESTIONS FROM TEXTBOOK
π΅ Question 1. Draw the diagram of a sarcomere of skeletal muscle showing different regions.
π’ Answer:
π§ A sarcomere is the functional unit of a myofibril, extending between two Z-lines.
𧬠Regions:
A-band: Dark band with both actin and myosin.
I-band: Light band with actin only.
H-zone: Central part of A-band with myosin only.
M-line: Middle line of H-zone.
Z-line: Boundary of sarcomere.
π‘ During contraction: I-band and H-zone shorten; A-band remains constant.
βοΈ (Diagram to be drawn showing Z-line, A-band, I-band, H-zone, M-line)
π΅ Question 2. Define sliding filament theory of muscle contraction.
π’ Answer:
βοΈ Sliding Filament Theory:
Explains muscle contraction due to sliding of thin (actin) filaments over thick (myosin) filaments.
Filaments do not change length, sarcomere shortens.
π‘ Powered by ATP hydrolysis and CaΒ²βΊ ions, forming cross-bridges between actin and myosin.
βοΈ Result: Shortening of muscle fibre β contraction.
π΅ Question 3. Describe the important steps in muscle contraction.
π’ Answer:
𧬠Steps:
Impulse from motor neuron β releases acetylcholine β depolarises sarcolemma.
CaΒ²βΊ ions released from sarcoplasmic reticulum.
CaΒ²βΊ binds troponin, shifting tropomyosin, exposing binding sites on actin.
Myosin head binds actin β cross-bridge forms.
ATP hydrolysis β power stroke β filaments slide.
ATP binding detaches myosin head β next cycle begins.
CaΒ²βΊ reabsorbed β relaxation.
βοΈ Requires ATP and CaΒ²βΊ.
π΅ Question 4. Write true or false. If false, change the statement so that it is true.
π’ Answer:
(a) Actin is present in thin filament β βοΈ True
(b) H-zone of striated muscle fibre represents both thick and thin filaments β β False
β‘οΈ Correct: H-zone has only thick (myosin) filaments.
(c) Human skeleton has 206 bones β βοΈ True
(d) There are 11 pairs of ribs in man β β False
β‘οΈ Correct: There are 12 pairs of ribs.
(e) Sternum is present on the ventral side of the body β βοΈ True
π΅ Question 5. Write the difference between:
π’ Answer:
(a) πΏ Actin and Myosin
Feature Actin Myosin
Type Thin filament Thick filament
Composition Actin, tropomyosin, troponin Myosin protein
Function Slides over myosin Cross-bridge formation
(b) πΈ Red and White muscles
Feature Red Muscle White Muscle
Myoglobin High Low
Mitochondria Many Few
Endurance High Low
Fatigue Resistant Quickly fatigued
(c) 𦴠Pectoral and Pelvic girdle
Feature Pectoral Pelvic
Function Supports forelimbs Supports hindlimbs
Bones Clavicle, scapula Ilium, ischium, pubis
Location Upper trunk Lower trunk
π΅ Question 6. Match Column I with Column II:
Column I Column II
(a) Smooth muscle (iv) Involuntary
(b) Tropomyosin (ii) Thin filament
(c) Red muscle (i) Myoglobin
(d) Skull (iii) Sutures
βοΈ Correct matches: (a)-(iv), (b)-(ii), (c)-(i), (d)-(iii)
π΅ Question 7. What are the different types of movements exhibited by the cells of human body?
π’ Answer:
πΏ Types:
Amoeboid movement β leucocytes, macrophages.
Ciliary movement β lining of trachea, fallopian tubes.
Muscular movement β skeletal muscles, limbs.
βοΈ All help in locomotion and internal transport.
π΅ Question 8. How do you distinguish between a skeletal muscle and a cardiac muscle?
π’ Answer:
Feature Skeletal Muscle Cardiac Muscle
Control Voluntary Involuntary
Nucleus Multinucleated Single nucleus
Intercalated discs Absent Present
Fatigue Rapid Never fatigues
Function Body movement Pumping of blood
π΅ Question 9. Name the type of joint between the following:
π’ Answer:
(a) Atlas/Axis β Pivot joint
(b) Carpal/Metacarpal of thumb β Saddle joint
(c) Between phalanges β Hinge joint
(d) Femur/Acetabulum β Ball and socket joint
(e) Between cranial bones β Sutures (fibrous joint)
(f) Between pubic bones in pelvic girdle β Cartilaginous joint
π΅ Question 10. Fill in the blank spaces:
π’ Answer:
(a) All mammals (except a few) have 7 cervical vertebrae.
(b) The number of phalanges in each limb of human is 14.
(c) Thin filament of myofibril contains 2 ‘F’ actins and two other proteins namely tropomyosin and troponin.
(d) In a muscle fibre, CaΒ²βΊ is stored in sarcoplasmic reticulum.
(e) 11th and 12th pairs of ribs are called floating ribs.
(f) The human cranium is made of 8 bones.
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OTHER IMPORTANT QUESTIONS FOR EXAMS
(CBSE MODEL QUESTIONS PAPER)
ESPECIALLY MADE FROM THIS LESSON ONLY
π΄ Question 1:
Which protein is responsible for muscle contraction?
π΄1οΈβ£ Keratin
π’2οΈβ£ Actin and Myosin
π‘3οΈβ£ Collagen
π΅4οΈβ£ Elastin
π’ Answer: 2οΈβ£ Actin and Myosin
π΄ Question 2:
Functional unit of muscle contraction is β
π΄1οΈβ£ Myofibril
π’2οΈβ£ Sarcomere
π‘3οΈβ£ Muscle fibre
π΅4οΈβ£ Fascicle
π’ Answer: 2οΈβ£ Sarcomere
π΄ Question 3:
Which of the following shows amoeboid movement in humans?
π΄1οΈβ£ Muscle cells
π’2οΈβ£ Leucocytes
π‘3οΈβ£ RBCs
π΅4οΈβ£ Neurons
π’ Answer: 2οΈβ£ Leucocytes
π΄ Question 4:
Which joint allows movement in all directions?
π΄1οΈβ£ Hinge joint
π’2οΈβ£ Ball and socket joint
π‘3οΈβ£ Pivot joint
π΅4οΈβ£ Saddle joint
π’ Answer: 2οΈβ£ Ball and socket joint
π΄ Question 5:
Which part of the myofibril disappears during contraction?
π΄1οΈβ£ A band
π’2οΈβ£ H zone
π‘3οΈβ£ I band
π΅4οΈβ£ Z line
π’ Answer: 2οΈβ£ H zone
π΄ Question 6:
Which ion triggers muscle contraction?
π΄1οΈβ£ NaβΊ
π’2οΈβ£ CaΒ²βΊ
π‘3οΈβ£ KβΊ
π΅4οΈβ£ MgΒ²βΊ
π’ Answer: 2οΈβ£ CaΒ²βΊ
π΄ Question 7:
The longest bone in human body is β
π΄1οΈβ£ Humerus
π’2οΈβ£ Femur
π‘3οΈβ£ Tibia
π΅4οΈβ£ Radius
π’ Answer: 2οΈβ£ Femur
π΄ Question 8:
Which connective tissue joins muscle to bone?
π΄1οΈβ£ Ligament
π’2οΈβ£ Tendon
π‘3οΈβ£ Cartilage
π΅4οΈβ£ Fascia
π’ Answer: 2οΈβ£ Tendon
π΄ Question 9:
The number of bones in adult human skeleton is β
π΄1οΈβ£ 200
π’2οΈβ£ 206
π‘3οΈβ£ 210
π΅4οΈβ£ 208
π’ Answer: 2οΈβ£ 206
π΄ Question 10:
Which enzyme hydrolyses ATP during muscle contraction?
π΄1οΈβ£ ATP synthase
π’2οΈβ£ Myosin ATPase
π‘3οΈβ£ Actinase
π΅4οΈβ£ Kinase
π’ Answer: 2οΈβ£ Myosin ATPase
π΄ Question 11:
Define locomotion.
π’ Answer:
The voluntary movement of an organism from one place to another is called locomotion (e.g., walking, running).
π΄ Question 12:
Name the three types of muscles in the human body.
π’ Answer:
1οΈβ£ Skeletal (Striated) β Voluntary, multinucleated
2οΈβ£ Smooth (Unstriated) β Involuntary, single nucleus
3οΈβ£ Cardiac β Involuntary, striated, branched
π΄ Question 13:
Define movement and locomotion. Differentiate between them.
π’ Answer:
Movement: Change in position of a body part (can be voluntary or involuntary).
Locomotion: Voluntary movement of the entire organism from one place to another.
Feature Movement Locomotion
Scope May be of a part Whole body
Voluntary Can be voluntary/involuntary Always voluntary
Example Beating of heart Walking, running
π‘ All locomotions are movements, but all movements are not locomotions.
π΄ Question 14:
What are the different types of movements in animals?
π’ Answer:
1οΈβ£ Amoeboid movement:
ββ In leucocytes; uses pseudopodia.
2οΈβ£ Ciliary movement:
ββ In respiratory tract & fallopian tubes; uses cilia.
3οΈβ£ Muscular movement:
ββ Due to muscle contraction; e.g., walking, running.
π‘ Human body shows all three types of movement.
π΄ Question 15:
Explain the structure of a sarcomere.
π’ Answer:
Definition: Structural & functional unit of myofibril (between two Z-lines).
Components:
β1οΈβ£ A band (dark): Contains thick filament (myosin).
β2οΈβ£ I band (light): Contains thin filament (actin).
β3οΈβ£ H zone: Central part with only myosin.
β4οΈβ£ Z line: Boundary of sarcomere.
π‘ During contraction, H zone shortens, A band remains same.
π΄ Question 16:
Describe the chemical events of muscle contraction.
π’ Answer:
1οΈβ£ Nerve impulse β releases CaΒ²βΊ from sarcoplasmic reticulum.
2οΈβ£ CaΒ²βΊ binds to troponin, exposing binding sites on actin.
3οΈβ£ Myosin head binds to actin β forms cross bridge.
4οΈβ£ ATP hydrolysis by myosin ATPase β energy for sliding.
5οΈβ£ Actin filaments slide β sarcomere shortens.
β
Relaxation when CaΒ²βΊ returns to SR.
π΄ Question 17:
List the functions of skeleton.
π’ Answer:
1οΈβ£ Support: Framework maintaining body shape.
2οΈβ£ Protection: Vital organs (skullβbrain, ribsβheart).
3οΈβ£ Movement: Bones act as levers with muscles.
4οΈβ£ Haemopoiesis: Bone marrow forms blood cells.
5οΈβ£ Mineral storage: Ca, P.
π‘ Total 206 bones in adult human.
π΄ Question 18:
Differentiate between axial and appendicular skeleton.
π’ Answer:
Feature Axial Appendicular
Components Skull, vertebral column, ribs, sternum Limbs & girdles
Bones 80 126
Function Protection & support Movement
Example Ribs Humerus
π‘ Together form complete skeleton.
π΄ Question 19:
What are synovial joints? Name their types.
π’ Answer:
Definition: Freely movable joints with synovial fluid.
Types:
β1οΈβ£ Ball & Socket β Shoulder, hip.
β2οΈβ£ Hinge β Elbow, knee.
β3οΈβ£ Pivot β Atlas-axis joint.
β4οΈβ£ Gliding β Carpals.
β5οΈβ£ Saddle β Thumb joint.
π‘ Enable various types of movements.
π΄ Question 20:
Write the differences between red and white muscles.
π’ Answer:
Feature Red Muscles White Muscles
Myoglobin High Low
Mitochondria Many Few
Endurance High Low
Colour Dark red Pale
Function Sustained activity Quick action
π‘ Red = aerobic; White = anaerobic.
π΄ Question 21:
Describe the structure of a typical vertebra.
π’ Answer:
Components:
β1οΈβ£ Centrum: Main body.
β2οΈβ£ Neural canal: For spinal cord.
β3οΈβ£ Processes: Transverse and spinous for muscle attachment.
Types: Cervical (7), Thoracic (12), Lumbar (5).
π‘ Vertebral column provides support & protection.
π΄ Question 22:
List the three types of muscles with their properties.
π’ Answer:
Type Control Striation Nucleus Function
Skeletal Voluntary Striated Multi Locomotion
Smooth Involuntary Unstriated Single Internal organs
Cardiac Involuntary Striated Single Heartbeat
π‘ Cardiac muscles are myogenic & fatigue-resistant.
π΄ Question 23:
Explain the sliding filament theory of muscle contraction.
π’ Answer:
Concept: Muscle contraction occurs due to sliding of thin (actin) filaments over thick (myosin) filaments.
Steps:
β1οΈβ£ Impulse: Nerve impulse releases CaΒ²βΊ from sarcoplasmic reticulum.
β2οΈβ£ Exposure: CaΒ²βΊ binds to troponin, shifts tropomyosin, exposes binding sites on actin.
β3οΈβ£ Cross-bridge: Myosin heads attach to actin β form cross-bridges.
β4οΈβ£ Power stroke: Myosin heads use ATP to pull actin inward β sarcomere shortens.
β5οΈβ£ Detachment: ATP binds again β myosin detaches; cycle repeats.
π‘ A band constant, I band & H zone shorten β muscle contracts.
π΄ Question 24:
Describe the structure of human skeleton.
π’ Answer:
Divisions:
β1οΈβ£ Axial skeleton (80 bones): Skull (22), vertebral column (26), ribs (24), sternum (1), hyoid (1), ear ossicles (6).
β2οΈβ£ Appendicular skeleton (126 bones): Limbs & girdles (pectoral and pelvic).
Functions:
ββοΈ Support and protection of organs.
ββοΈ Locomotion with muscles.
ββοΈ Haemopoiesis in bone marrow.
ββοΈ Mineral storage (Ca, P).
π‘ Total = 206 bones in adult human.
π΄ Question 25:
Explain the structure of a typical myofibril.
π’ Answer:
Myofibril: Cylindrical structure inside muscle fibre.
Bands:
β1οΈβ£ A band (dark): Thick myosin filaments.
β2οΈβ£ I band (light): Thin actin filaments.
β3οΈβ£ H zone: Central part with only myosin.
β4οΈβ£ Z line: Border between sarcomeres.
Sarcomere: Segment between two Z-lines = functional unit.
π‘ Contraction: actin slides over myosin β sarcomere shortens.
π΄ Question 26:
Describe the events of muscle contraction and relaxation.
π’ Answer:
Contraction:
1οΈβ£ Impulse from motor neuron β CaΒ²βΊ released.
2οΈβ£ CaΒ²βΊ binds troponin β exposes actin sites.
3οΈβ£ Myosin binds actin β cross-bridge forms.
4οΈβ£ ATP hydrolysis β power stroke β filaments slide.
Relaxation:
1οΈβ£ CaΒ²βΊ pumped back to sarcoplasmic reticulum.
2οΈβ£ Tropomyosin covers actin sites.
3οΈβ£ Cross-bridges detach β muscle returns to rest.
π‘ Energy: Provided by ATP; lack of ATP β rigor mortis.
π΄ Question 27:
Explain the structure and functions of synovial joint.
π’ Answer:
Structure:
ββ Ends of bones covered with cartilage.
ββ Enclosed by joint capsule lined with synovial membrane.
ββ Synovial fluid lubricates and reduces friction.
Functions:
ββοΈ Free movement in various directions.
ββοΈ Shock absorption.
ββοΈ Stability and flexibility.
Examples: Shoulder (ball & socket), elbow (hinge).
π΄ Question 28:
Describe the disorders of the muscular and skeletal system.
π’ Answer:
1οΈβ£ Myasthenia gravis: Autoimmune; weak neuromuscular transmission.
2οΈβ£ Tetany: Low CaΒ²βΊ β spasms.
3οΈβ£ Muscular dystrophy: Degeneration of muscles.
4οΈβ£ Arthritis: Inflammation of joints.
5οΈβ£ Osteoporosis: Bone weakening due to low calcium.
π‘ Balanced diet & exercise prevent many disorders.
π΄ Question 29:
Explain the types and functions of joints in the human body.
π’ Answer:
1οΈβ£ Fibrous joints: Immovable (e.g., skull sutures).
2οΈβ£ Cartilaginous joints: Slightly movable (e.g., vertebrae).
3οΈβ£ Synovial joints: Freely movable; types:
ββ Ball & socket (hip)
ββ Hinge (elbow)
ββ Pivot (neck)
ββ Gliding (carpals)
ββ Saddle (thumb)
π‘ Functions: Movement, flexibility, shock absorption.
π΄ Question 30:
Describe the appendicular skeleton and its role.
π’ Answer:
Components (126 bones):
β1οΈβ£ Pectoral girdle (4): Clavicle & scapula.
β2οΈβ£ Pelvic girdle (2): Hip bones.
β3οΈβ£ Limbs (120):
βββ Upper limb: Humerus, radius, ulna, carpals, metacarpals, phalanges.
βββ Lower limb: Femur, tibia, fibula, tarsals, metatarsals, phalanges.
Functions:
ββοΈ Locomotion and manipulation.
ββοΈ Attachment of muscles.
ββοΈ Protection (pelvis β reproductive organs).
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